LOS ANGELES—After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

The California zoo invited fans of the late Galápagos tortoise to enjoy one of her favorite snacks, a "generous, fruit-filled salad," in her honor San Diego Zoo/I/nstagram Gramma, the Galápagos ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

Gramma, one of the San Diego Zoo’s longest-living Galápagos tortoises, died on November 20, 2025, at the estimated age of 141, the institution confirmed. The veterinary team performed humane ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Woman's Late ...

SAN DIEGO — One of the first animals brought to the San Diego Zoo has died at the age of 141 years old. The zoo announced the death of its beloved Galápagos tortoise, Gramma, on social media, saying ...